Summary
In a large random mating population segregating for a pair of allelomorphs with full dominance, the condition that neither allelomorph should disappear is that the arithmetic mean of the fitnesses of recessives in different generations should exceed unity, their geometric mean fall below it.
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References
Haldane, J. B. S. (1924). A mathematical theory of natural and artificial selection, Part I.Trans. Camb. Phil. Soc.,23, 19–41.
Fisher, R. A. (1922). On the dominance ratio.Proc. Roy. Soc. Edin.,42, 321–341.
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Haldane, J.B.S., Jayakar, S.D. Polymorphism due to selection of varying direction. J Genet 58, 237–242 (1963). https://doi.org/10.1007/BF02986143
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DOI: https://doi.org/10.1007/BF02986143